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Body size and metabolic rate
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It is much easier for a small animal to run uphill than for a large animal, because
Smaller animals have higher mass-specific metabolic rates.
The metabolic rate of an organism at rest at normal temperature is its basal metabolic rate (BMR). BMR is inversely proportional to body weight.
Small animals have a higher BMR, so they get more energy and can run uphill more easily than larger animals.